…the main mode of human adaptation is social learning mechanisms that operate over multiple generations.
…the behavior of a certain tribe, whether in constructing baskets or following certain marriage practices, is largely due to the fact that their ancestors hundreds or even thousands of years ago practiced the behavior. This means that there is considerable cultural inertia in human behavior, which may have persisted for up to 15,000 years.
My comment: It means the behavior is nutrient-dependent and RNA-mediated via the fixation of amino acid substitutions in the context of the physiology of reproduction. That fact links the epigenetic landscape to the physical landscape of DNA via metabolic and genetic networks in all genera.
See for examples of how amino acid substitutions are linked from biophysically constrained protein folding to biodiversity: